So basically, somebody simply re-barrelled the 410 smooth bore musket with a 303 rifled barrel and then cancelled the 410 marking?
This would have been done post 1942 (410 conversion date). Well, there was a war going on at that time. Somebody needed a rifle more than a shotgun?
Maybe we can glean some clues and determine who did this if we could see the markings on the barrel reinforce either side of the Nock's form and the underside.