OK just for closure. So it seems the .303 bolt that was supposed to be sent back to me when I converted from 54R was NOT a .303 bolt but the same 54R bolt as before. So... the spent casing was just rattling around on the face of the breach block and was getting dropped into the receiver during cycling and jamming it up. I replaced the other block with a (correct) .303 block and it ran perfectly without missing a beat. Thanks to all to helped me trouble shoot this. I very much appreciate all your time and effort.
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