Take the time to read Peter's articles listed on this site about working these rifles.

Greek and other European returns don't necessarily have any foreign markings as i said and not all Greek rifles had the magazines numbered although the majority do. The magazine could have also been replaced.

I make the observation having inspected several hundred Long Branch No.4's over many years. The font is different than what it left the factory with, (look at the serial number on the butt socket), and the rear side of the bolt body looks like it's been filed and renumbered. This is common and correct for a rifle that's been worked on in service. Is it a Long Branch bolt body? I can't really tell from your pictures. Your rifle would have likely had a Mk.2 flip sight as original so it's been through workshops a time or two for sure. I still sense a European return. It's a nice, honest rifle IMHO.

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I'm not sure what the "O" and "G2" are. Guessing they are inspectors stamps used during production.