Danny, I just knocked up a rough sketch in paint to show how the o ring will centre the base of the case in the back of the chamber.
The rubber o ring is uniform is section all the way around and flexible. It will be squeezed into the space evenly all the way around by the bolt being closed onto the case. This should centre the rear of the cartridge regardless of if the boltface is square to the chamber or not.
I think the o ring would be firm enough though to negate most of the effect of the extractor bearing on the right side. In fact, if you watch the extractor closely as you cam the bolt shut, you may see the extractor being pushed outward by the case being centred- either that or it's seating onto the machined slot on the barrel- taking the weight off the case.