A lot of "target" rifles were handed back "for the duration" in WW2
Some of these ended up with the VDC, (Volunteer Defence Corps). A lot of members of this organisation already had a rifle (or two) in .303. I notice that the one in the AWM pic still has its magazine cutoff, despite these being officially "withdrawn" TWICE. First time was with the introduction of the No1 Mklll* "economy model. Post WW1, it was decided to put the fruit back on. The second time "economies" were introduced again was not long before WW2.